1. Department of Biosciences and Informatics, Faculty of Science and Technology, Keio University, Kohoku, Yokohama 223-8522, Japan.
2. Institute for Integrative Neurobiology and Department of Biology, Konan University, Kobe 658-8501, Japan.
3. Shimoda Marine Research Center, University of Tsukuba, Shimoda 415-0025, Japan.