1. Marine Spatial Ecology Lab, School of Biological Sciences, University of Queensland, St Lucia, Queensland 4072, Australia.
2. Australian Research Council Centre of Excellence for Coral Reef Studies, University of Queensland, St Lucia, Queensland 4072, Australia.
3. Global Science, The Nature Conservancy, Brunswick, ME 04011, USA.
4. Australian Institute of Marine Science, PMB3, Townsville, Queensland 4810, Australia.
5. Centre for Environment, Fisheries and Aquaculture Science, Lowestoft NR33 0HT, UK.
6. Catchment to Reef Research Group, Centre for Tropical Water and Aquatic Ecosystem Research, James Cook University, Townsville, Queensland 4811, Australia.