1. Key Laboratory of Vertebrate Evolution and Human Origins, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, 142 Xi Zhi Men Wai Street, Beijing 100044, China.
2. Division of Paleontology and Richard Gilder Graduate School, American Museum of Natural History, Central Park West at 79th Street, New York, NY 10024, USA.
3. CAS Center for Excellence in Life and Paleoenvironment, Beijing 100044, China.
4. University of Chinese Academy of Sciences, Beijing 100049, China.
5. Department of Earth Science, University of California, Santa Barbara, Santa Barbara, CA 93106, USA.