1. Department of Comparative Medicine, Yale School of Medicine, New Haven, CT 06519, USA.
2. Department of Immunobiology, Yale School of Medicine, New Haven, CT 06519, USA.
3. Novartis Institutes for BioMedical Research, 250 Massachusetts Ave, Cambridge, MA 02139, USA.
4. Department of Molecular Biophysics and Biochemistry, W.M. Keck Foundation Biotechnology Resource Laboratory, Yale University School of Medicine, New Haven, CT 06520, USA.
5. Yale Center for Research on Aging, Yale School of Medicine, New Haven, CT 06519, USA.
6. Howard Hughes Medical Institute, Chevy Chase, MD 20815, USA.
7. Department of Molecular Cellular and Developmental Biology, Yale University, New Haven, CT, 06511, USA.