1. Department of Cell and Developmental Biology, University College London (UCL), Gower Street, London WC1E 6BT, UK.
2. School of Life, Health and Chemical Sciences, Faculty of Science, Technology, Engineering and Mathematics, Walton Hall, Open University, Milton Keynes MK7 6AA, UK.
3. Department of Pharmacology, University of Cambridge, Tennis Court Road, Cambridge CB2 1PD, UK.
4. Department of Cell Biology, Neurobiology and Anatomy, Medical College of Wisconsin, Milwaukee WI 53226, USA.