1. Department of Molecular, Cellular and Developmental Biology, University of Michigan, Ann Arbor, MI 48109, USA.
2. The Key Laboratory of Mariculture, Education Ministry of China and College of Fisheries, Ocean University of China, Qingdao 266003, China.
3. Pilot National Laboratory for Marine Science and Technology (Qingdao), Qingdao 266237, China.