1. Australian Research Council Centre of Excellence for Coral Reef Studies, University of Western Australia, Crawley, Western Australia 6009, Australia.
2. School of Earth and Environment, University of Western Australia, Crawley, Western Australia 6009, Australia.
3. University of Western Australia Oceans Institute, University of Western Australia, Crawley, Western Australia 6009, Australia.
4. Western Australian Marine Science Institution, Floreat, Western Australia 6014, Australia.