1. Department of Medicine, University of North Carolina at Chapel Hill, Chapel Hill, NC 27599, USA.
2. Department of Medicine, University of Alabama at Birmingham, Birmingham, AL 35294, USA.
3. Department of Cell Biology and Physiology, and Lineberger Comprehensive Cancer Center, University of North Carolina at Chapel Hill, Chapel Hill, NC 27599, USA.
4. Department of Pathology, University of Alabama at Birmingham, Birmingham, AL 35294, USA.
5. Program in Protease and Matrix Biology, University of Alabama at Birmingham, Birmingham, AL 35294, USA.
6. Faculty of Science, Utrecht Institute for Pharmaceutical Sciences, Utrecht University, 3508 TB Utrecht, Netherlands.
7. Lung Health Center, University of Alabama at Birmingham, Birmingham, AL 35294, USA.
8. Medical Service at Birmingham VA Medical Center, Birmingham, AL 35233, USA.
9. Department of Cell, Developmental, and Integrative Biology, University of Alabama at Birmingham, Birmingham, AL 35294, USA.