1. Department of Life, Earth, and Environmental Sciences, West Texas A&M University, Canyon, TX 79016, USA.
2. US Fish and Wildlife Service, Desert Tortoise Recovery Office, Reno, NV 89502, USA.
3. Department of Ecology and Evolutionary Biology, University of California, Los Angeles, CA 90095, USA.
4. La Kretz Center for California Conservation Science, Institute of the Environment and Sustainability, University of California, Los Angeles, CA 90095, USA.