1. Centre for Biological Diversity and Scottish Oceans Institute, School of Biology, University of St. Andrews, St. Andrews, Fife KY16 9TH, UK.
2. Department of Biology, University of Vermont, Burlington, VT 05405, USA.
3. School of Biology and Ecology, Sustainability Solutions Initiative, University of Maine, Orono, ME 04469, USA.
4. Department of Mathematics, Keio University, 3-14-1 Hiyoshi Kohoku, Yokohama 223-8522, Japan.