1. State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, 650223 Kunming, China.
2. Kunming College of Life Science, University of Chinese Academy of Sciences, 650204 Kunming, China.
3. Centre for Biodiversity, Royal Ontario Museum, Toronto, ON M5S 2C6, Canada.
4. Center for Excellence in Animal Evolution and Genetics, Chinese Academy of Sciences, 650223 Kunming, China.