1. Australian Research Council (ARC) Centre of Excellence for Coral Reef Studies, St. Lucia, Queensland 4072, Australia.
2. School of Biological Sciences, University of Queensland, St. Lucia, Queensland 4072, Australia.
3. ARC Centre of Excellence for Coral Reef Studies and School of Marine and Tropical Biology, James Cook University, Townsville, Queensland 4011, Australia.
4. Department of Geology and Geophysics, Woods Hole Oceanographic Institution, Woods Hole, MA 02543, USA.