1. Department of Biology, 108 Coker Hall, University of North Carolina (UNC), CB 3280, Chapel Hill, NC 27599–3280, USA.
2. VIB Department of Plant Systems Biology and Department of Plant Biotechnology and Genetics, Ghent University, 9052 Ghent, Belgium.
3. Curriculum in Genetics and Molecular Biology and Department of Microbiology and Immunology, UNC, Chapel Hill, NC 27599, USA.
4. Carolina Center for Genome Sciences, UNC, Chapel Hill, NC 27599, USA.