1. Department of Zoology, University of Oxford, South Parks Road, Oxford, OX1 3PS, UK.
2. Centre for Ecology and Hydrology, Maclean Building, Benson Lane, Crowmarsh Gifford, Wallingford, Oxfordshire, OX10 8BB, UK.
3. Centre for Conservation Ecology and Environmental Change, School of Conservation Sciences, Bournemouth University, Fern Barrow, Talbot Campus, Poole, Dorset BH12 5BB, UK.