1. School of Oceanography, University of Washington, Seattle, WA 98195, USA.
2. Department of Atmospheric and Oceanic Sciences, University of California, Los Angeles, CA 90095, USA.
3. Biological Sciences Department, University of Rhode Island, Kingston, RI 02881, USA.
4. Alfred Wegener Institute, D-27570 Bremerhaven, Germany.
5. Department of Biology, University of Washington, Seattle, WA 98195, USA.