1. Max Planck Institute for Physics of Complex Systems, Nöthnitzer Straße 38, 01187 Dresden, Germany.
2. Department of Ecology and Evolutionary Biology and Center for BioComplexity, Princeton University, Princeton, NJ 08540, USA.
3. University Fellow, Resources for the Future, 1616 P Street NW, Washington, DC 20036, USA.
4. Evolution and Ecology Program, International Institute for Applied Systems Analysis, Schlossplatz 1, 2361 Laxenburg, Austria.