1. Sainsbury Laboratory, University of Cambridge, Bateman Street, Cambridge CB2 1LR, UK.
2. Department of Applied Mathematics and Theoretical Physics, University of Cambridge, Cambridge CB3 0WA, UK.
3. Department of Astronomy and Theoretical Physics, Lund University, SE22362 Lund, Sweden.
4. Howard Hughes Medical Institute and Division of Biology and Biological Engineering, California Institute of Technology, 1200 East California Boulevard, Pasadena, CA 91125, USA.