1. Department of Biological Sciences, SAF Building, Imperial College London, South Kensington Campus, London SW7 2AZ, UK.
2. NERC Centre for Population Biology, Imperial College London, Silwood Park Campus, Ascot, Berkshire SL5 7PY, UK.
3. University of Perugia, Via del Giochetto, 06100 Perugia, Italy.