1. Department of Ecology and Evolutionary Biology, University of Toronto, Toronto, Ontario M5S 3B2, Canada.
2. Department of Biology, McGill University, Montreal, Quebec H3A 1B1, Canada.
3. Institut de Genomique Fonctionnelle de Lyon, Ecole Normale Supérieure, CNRS UMR 5242, 46 allée d’Italie, 69364 Lyon Cedex 07, France.