1. Department of Anatomy and Brain Science, Kobe University School of Medicine, Kobe 650-0017, Japan.
2. Department of Molecular Genetics, Institute of Development, Aging and Cancer, Tohoku University, Sendai 980-8575, Japan.
3. MGC, Department of Cell Biology and Genetics, Erasmus University, Post Office Box 1738, 3000 DR Rotterdam, Netherlands.