1. P. M. Vitousek and H. A. Mooney are in the Department of Biological Sciences, Stanford University, Stanford, CA 94305, USA. J. Lubchenco is in the Department of Zoology, Oregon State University, Corvallis, OR 97331, USA. J. M. Melillo is at the U.S. Office of Science and Technology Policy, Old Executive Office Building, Room 443, Washington, DC 20502, USA.