1. Graduate School of Agriculture, Kyoto University, Kyoto, Japan.
2. Graduate School of Bioscience and Bioindustry, Tokushima University, Tokushima, Japan.
3. Graduate School of Biological Sciences, Nara Institute of Science and Technology, Ikoma, Japan.
4. Japan Science and Technology Agency (JST), Precursory Research for Embryonic Science and Technology (PRESTO), Kawaguchi, Japan.
5. RIKEN Center for Sustainable Resource Science, Yokohama, Japan.