1. Global Ocean Team, The Nature Conservancy, Department of Physical, Earth and Environmental Sciences, University of Siena, Pian dei Mantellini, 44, 53100 Siena, Italy; and Department of Zoology, University of Cambridge, Cambridge CB2 3EJ, UK.
2. School of Biology, Newcastle University, Newcastle NE1 7RU, UK; and Environmental Research Institute, University of the Highlands and Islands, Thurso KW14 7EE, UK.