1. Department of Oceanography, Dalhousie University, Halifax, Nova Scotia, B3H 4J1 Canada.
2. Department of Fisheries and Oceans, Marine Ecology Laboratory, Bedford Institute of Oceanography, Dartmouth, Nova Scotia, B2Y 4A2 Canada.
3. Department of Fisheries and Oceans, Atlantic Oceanographic Laboratory, Bedford Institute of Oceanography, Dartmouth, Nova Scotia, B2Y 4A2 Canada.