1. Department of Ecology, Evolution, and Marine Biology, University of California, Santa Barbara, CA 93106, USA.
2. Department of Ecology, Evolution, and Natural Resources, Institute of Marine and Coastal Sciences, Rutgers University, New Brunswick, NJ 08901, USA.
3. Department of Biology, Stanford University, Hopkins Marine Station, Pacific Grove, CA 93950, USA.
4. Department of Ecology and Evolutionary Biology, University of California, Santa Cruz, CA 95060, USA.