1. School of Life Sciences, Arizona State University, Tempe, AZ 85287–4501, USA.
2. New Zealand Centre for Ecological Economics, Private Bag 11 052, Palmerston North, New Zealand.
3. School of Sustainability, Arizona State University, Tempe, AZ 85287–3211, USA.
4. CSIRO Sustainable Ecosystems, Canberra ACT 2601, Australia.