1. Organismic and Evolutionary Biology Program, University of Massachusetts, Amherst, MA 01003–5810, USA.
2. Marine Molecular Biotechnology Laboratory, University of Washington, 3707 Brooklyn Avenue Northeast, Seattle, WA 98105–6715, USA.
3. School of Aquatic and Fishery Sciences, University of Washington, Box 355020, Seattle, WA 98195, USA.
4. Washington Department of Fish and Wildlife, 600 Capitol Way North, Olympia, WA 98501, USA.