1. Institute of Science and Technology Austria, 3400 Klosterneuburg, Austria.
2. Faculty of Arts and Sciences, Center for Systems Biology and Department of Molecular and Cellular Biology, Harvard University, Cambridge, MA 02138, USA.
3. Department of Environmental System Science, ETH Zurich, Swiss Federal Institute of Technology, 8092 Zurich, Switzerland, and Department of Environmental Microbiology, Eawag, Swiss Federal Institute of Aquatic Science and Technology, 8600 Duebendorf, Switzerland.