1. Laboratory of Persistent Viral Diseases, Rocky Mountain Laboratories, National Institute of Allergy and Infectious Diseases, Hamilton, MT 59840, USA.
2. Division of Virology, Department of Neuropharmacology, Scripps Research Institute, La Jolla, CA 92037, USA.
3. Departments of Neurosciences and Pathology, University of California, San Diego, La Jolla, CA 92093, USA.