1. Department of Cell Biology, Harvard Medical School, Boston, MA 02115, USA.
2. Department of Pathology, Boston Children’s Hospital and Harvard Medical School, Boston, MA 02115, USA.
3. Department of Hematology, St. Jude Children’s Research Hospital, Memphis, TN 38105, USA.
4. Genetics and Genome Biology Program, Hospital for Sick Children, Peter Gilgan Centre for Research and Learning, Toronto, Ontario M5G 0A4, Canada.