1. Key Laboratory of Vertebrate Evolution and Human Origins, Institute of Vertebrate Paleontology and Paleoanthropology (IVPP), Chinese Academy of Sciences, 142 Xi Zhi Men Wai Street, Beijing, 100044, China.
2. Chinese Academy of Sciences (CAS) Center for Excellence in Tibetan Plateau Earth Sciences, Beijing, 100101, China.
3. Biodiversity Institute, University of Kansas, 1345 Jayhawk Boulevard, Lawrence, KS 66045-7561, USA.
4. Department of Ecology and Evolutionary Biology, University of Kansas, 1345 Jayhawk Boulevard, Lawrence, KS 66045-7561, USA.