1. Department of Animal and Plant Sciences, University of Sheffield, Sheffield S10 2TN, UK.
2. The Marine Biological Association of the United Kingdom, Citadel Hill, Plymouth PL1 2PB, UK.
3. Department of Plant and Environmental Protection Sciences, University of Hawaii at Manoa, Hawaii, USA.
4. The Food and Environment Research Agency, Sand Hutton, York YO41 1LZ, UK.