1. Co-Innovation Center for Sustainable Forestry in Southern China & Key Laboratory of National Forestry and Grassland Administration on Subtropical Forest Biodiversity Conservation, College of Biology and the Environment, Nanjing Forestry University, Nanjing 210037, China.
2. Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming 650201, China.
3. Missouri Botanical Garden, St. Louis, MO 63110, USA.
4. Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, China.