1. Molecular Microbial Ecology Section, Natural Environment Research Council (NERC) Centre for Ecology and Hydrology, Mansfield Road, Oxford, OX1 3SR, UK.
2. Environmental Biotechnology Section, Natural Environment Research Council (NERC) Centre for Ecology and Hydrology, Mansfield Road, Oxford, OX1 3SR, UK.
3. Department of Zoology, University of Oxford, South Parks Road, Oxford, OX1 3PS, UK.
4. Department of Engineering Science, University of Oxford, Parks Road, Oxford, OX1 3PJ, UK.