1. Guangdong Basic Research Center of Excellence for Ecological Security and Green Development, Southern Marine Science and Engineering Guangdong Laboratory (Guangzhou), Guangzhou 511458, China.
2. Research Centre of Ecology & Environment for Coastal Area and Deep Sea, Southern Marine Science and Engineering Guangdong Laboratory (Guangzhou), Guangzhou 511458, China.
3. Guangdong Basic Research Center of Excellence for Ecological Security and Green Development, School of Ecology, Environment and Resources, Guangdong University of Technology, Guangzhou 510006, China.
4. Simon F. S. Li Marine Science Laboratory and School of Life Sciences, The Chinese University of Hong Kong, Shatin, Hong Kong SAR, China.