1. Whitehead Institute for Biomedical Research and Department of Biology, Massachusetts Institute of Technology, Cambridge, MA 02142, USA.
2. Ludwig Maximilians University, Department of Biology II, 80336 Munich, Germany.
3. Max Delbrück Center for Molecular Medicine, 13125 Berlin, Germany.
4. Department of Laboratory Medicine and UCSF Comprehensive Cancer Center, University of California, 2340 Sutter Street, San Francisco, CA 94143, USA.