1. Department of Plant Breeding and Genetics, Cornell University, Ithaca, NY 14853, USA.
2. Department of Plant Biology, Cornell University, Ithaca, NY 14853, USA.
3. Department of Plant Sciences, Arizona Genomics Institute, University of Arizona, Tucson, AZ 85721, USA.
4. U.S. Department of Agriculture, Agricultural Research Service, Plant, Soil, Nutrition Lab and Boyce Thompson Institute for Plant Research, Cornell University, Ithaca, NY 14853, USA.