1. Division of Neuroscience, Children's Hospital, Harvard Medical School, Boston, MA 02115, USA.
2. Program in Biological and Biomedical Sciences, Harvard Medical School, Boston, MA 02115, USA.
3. Whitehead Institute for Biomedical Research, Massachusetts Institute of Technology, Cambridge, MA 02142, USA.
4. Department of Biology, Massachusetts Institute of Technology, Cambridge, MA 02142, USA.