1. U.S. Department of Agriculture–Agricultural Research Service National Center for Genetic Resources Preservation, Fort Collins, CO 80521, USA.
2. Plant Germplasm Conservation Research, School of Life Sciences, University of KwaZulu-Natal, Durban, 4001 South Africa.
3. The Center for Plant Conservation, St. Louis, MO 63166-0299, USA.
4. Missouri Botanical Garden, St. Louis, MO 63166-0299, USA.