1. Department of Global Ecology, Carnegie Institution of Washington, Stanford, CA 94305, USA.
2. U.S. Department of Agriculture Forest Service, International Institute of Tropical Forestry, Rio Piedras, PR 100745, USA.
3. Complex Systems Research Center, Morse Hall, University of New Hampshire, Durham, NH 03824, USA.
4. Empresa Brasileira de Pesquisa Agropecuária-Amazonia Oriental, Trav. Dr Eneas Pinheiro SN, Belem CEP 66095–100, Pará, Brazil.