1. Key Laboratory of Vertebrate Evolution and Human Origins, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing 100044, China.
2. CAS Center for Excellence in Life and Paleoenvironment, Beijing 100044, China.
3. Division of Paleontology, American Museum of Natural History, New York, NY 10024, USA.
4. Microscopy and Imaging Facility, American Museum of Natural History, New York, NY 10024, USA.
5. Earth and Environmental Sciences, Graduate Center, City University of New York, New York, NY 10016, USA.