1. CAS Key Laboratory of Tropical Forest Ecology, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Mengla, Yunnan 666303, China.
2. College of Life Sciences, University of Chinese Academy of Sciences, Beijing 100049, China.
3. Negaunee Integrative Research Center, Field Museum, Chicago, IL 60605, USA.
4. Center of Plant Ecology, Core Botanical Gardens, Chinese Academy of Sciences, Mengla, Yunnan 666303, China.
5. School of Environment, Earth, and Ecosystem Sciences, The Open University, Milton Keynes MK7 6AA, UK.