1. Department of Grape and Persimmon Research, National Institute of Fruit Tree Science, Akitsu, Hiroshima 729-2494, Japan.
2. National Research Institute of Brewing, Higashi-Hiroshima, Hiroshima 739-0046, Japan.
3. Molecular Genetics Department, National Institute of Agrobiological Sciences, Tsukuba, Ibaraki 305-8602, Japan.