1. CEREGE, Collège de France, UMR 6635, CNRS Université Aix-Marseille III, Europole de l'Arbois, BP 80, 13545 Aix-en-Provence, France.
2. Department of Marine Biogeochemistry and Toxicology, Royal Netherlands Institute for Sea Research (NIOZ), Post Office Box 59, 1790 AB Den Burg, Texel, Netherlands.