1. S. J. McNaughton and M. M. McNaughton, Biological Research Laboratories, Syracuse University, Syracuse, NY 13244–1220, USA; and Serengeti Wildlife Research Centre, Post Office Box 661, Arusha, Tanzania.
2. F. F. Banyikwa, Department of Botany, University of Dar es Salaam, Post Office Box 35060, Dar es Salaam, Tanzania; Biological Research Laboratories, Syracuse University, Syracuse, NY 13244–1220, USA; and Serengeti Wildlife Research Centre, Post Office Box 661, Arusha, Tanzania.