1. University of Edinburgh Usher Institute, Edinburgh EH16 4UX, UK.
2. University of Edinburgh MRC Centre for Reproductive Health, Edinburgh EH16 4TJ, UK.
3. Wellcome Leap In Utero Program, Wellcome Leap Inc., Culver City, CA 90232, USA.
4. The Rosie Hospital and NIHR Cambridge Biomedical Research Centre, Department of Obstetrics and Gynaecology, University of Cambridge, Cambridge CB2 0SW, UK.
5. Centre for Trophoblast Research, Department of Obstetrics and Gynaecology, University of Cambridge, Cambridge CB2 0SW, UK.
6. Wellcome-MRC Institute of Metabolic Science and Medical Research Council Metabolic Diseases Unit, Addenbrooke’s Hospital, University of Cambridge, Cambridge CB2 0QQ, UK.