1. Center for Macroecology, Evolution and Climate, Globe Institute, University of Copenhagen, Copenhagen, Denmark.
2. Nature Uganda, Katalima Crescent Lower Naguru, Kampala, Uganda.
3. RSPB Centre for Conservation Science, The Lodge, Sandy, Bedfordshire SG19 2DL, UK.
4. Centre for Biodiversity and Environment Research, Department of Genetics, Evolution and Environment, University College London, London WC1E 6BT, UK.
5. Biodiversity, Ecology and Conservation Group, International Institute for Applied System Analyses, 2361 Laxenburg, Austria.
6. Nanjing Institute of Environmental Sciences, Ministry of Ecology and Environment of China, Nanjing 210042, China.