1. Institute of Marine and Coastal Sciences, Rutgers University, New Brunswick, NJ 08901, USA.
2. Department of Biology, Dalhousie University, 1355 Oxford Street, Halifax, NS B3H 4R2, Canada.
3. Centre For Ecological and Evolutionary Synthesis, Department of Biosciences, University of Oslo, Oslo, Norway.
4. Department of Biology, University of Victoria, Post Office Box 1700 STN CSC, Victoria, British Columbia V8W 2Y2, Canada.