1. College of Animal Sciences (College of Bee Science), Fujian Agriculture and Forestry University, Fuzhou, 350002, China.
2. Centre de Recherches sur la Cognition Animale, Centre de Biologie Intégrative (CBI), University of Toulouse, CNRS, UPS, 31062 Toulouse cedex 9, France.
3. Institut Universitaire de France (IUF), Paris, France.