1. Swedish University of Agricultural Sciences, Department of Aquatic Resources, Institute of Marine Research, Turistgatan 5, SE-453 30 Lysekil, Sweden.
2. Swedish Institute for the Marine Environment (SIME), Box 260, SE-405 30 Göteborg, Sweden.
3. Biology Department, Seattle University, 901 12th Avenue, Post Office Box 222000, Seattle, WA 98122-1090, USA.